Danser's Monograph on Nepenthes: Nepenthes Hookeriana



18. Nepenthes Hookeriana LINDL., Gard. Chron., 1848, p. 87 (1848) (non vidi) ; MAST., Gard. Chron., 1881, 2, p. 812, ic. 157 (1881) ; DIXON, Gard. Chron., 1888, 1, p. 170 (1888) ; MOTT., Dict., III, p. 449 (1896) ; ? BOERL., Handl., III, 1, p. 54 (1900) ; MACF., in ENGL., Pflanzenr., IV, 111, p. 34 (1908) ; in BAIL., Cycl., IV, p. 2127, ic. 4262, 4 (1919) ; MERR., Bibl. enum. Born., p. 283 (1921) ; non Low, Saraw., p. 68 (1848), cit. Gard. Chron., 1881, 2, p. 812, quae est N. Rafflesiana ; N. Loddigesii BAXT., in LOUD., Hort. Brit., suppl., III, p. 593 (1850) (non vidi) ; ? BECK, Wien. Ill. Gartenz., 1895, p. 277 (1895) ; BOERL., Handl., III, 1, p. 54 (1900) ; N. Rafflesiana, d) Hookeriana BECK, Wien. Ill. Gartenz., 1895, p. 147 (1895) ; N. hybrida g) elongata BECK, Wien. Ill. Gartenz., 1895, p. 221 (1895) (see. MACF.); N. elongata MACF., in ENGL., Pflanzenr., IV, 111, p. 81 (1908).

lcones: Gard. Chron., 1881, 2, p. 813 (1881), bona, asc. 1 inf.; Gard. Chron., 1892, 2, p. 557 (1882), idem ac praecedens ; Wien. Ill. Gartenz., 1895, p. 143, ic. 6 (1895), optima, asc. 1 inf.; BAIL., Cycl., IV, ic. 2462,4 (1919), optima, asc. 1 inf.; nostra 9.

Folia mediocria vix v. breviter petiolata, lamina lanceolato-spathulata, nervis longitudinalibus utrinque 4-5, vagina caulis c. 4/5 amplectente ; ascidia rosularum et inferiora oblique urceolata, alis 2 fimbriatis ; peristomio operculum versus in collum breve elongato, applanato, ad 10 mm lato, costis 1/2-1/3 mm distantibus, dentibus 3-5 x longioribus quam latis ; operculo elliptico-oblongo, facie inferiore plano ; ascidia superiora infundibuliformia, costis 2 prominentibus, raro alis 2 fimbriatis peristomio operculum versus in collum breve elongato, applanato, ad 10 mm lato, costis 1/2-1/3 mm distantibus, dentibus c. 3 x longioribus quam latis ; operculo elliptico-oblongo, facie inferiore plano ; inflorescentia racemus pedicellis inferioribus 12-15 mm longis, 2-floris,

. Fig. 9. Nepenthes Hookeriana ; a. upper portion of a stem with female flowers and fruits, 1/2 x (JAHERI 63) ; b. upper leaf with pitcher, 1/2 x (TEYSMANN 3512) ; c. rosette pitcher, 1/2 x (VILLAMIL 261 bis).

superioribus brevioribus 1-floris ; indumentum tomentosum e pilis stellatis intricatis compositum, denique maxima parte deciduum puncta fusca relinquens, in inflorescentiis permanens.

Stems climbing, the part with adult leaves cylindrical, 5 to 10 mm thick ; often short shoots and rosettes at the base of older plants. Leaves of the rosettes and the short shoots scattered, coriaceous, spathulate, 10 to 30 cm long, acute, gradually attenuate into a short, at most 8 cm long petiole, which forms a laterally flattened sheath, clasping the stem for about 4/5 ; pennate nerves numerous, running almost straightly towards the margin, the longitudinal ones 2 to 5 on each side, running parallel in the outer 2/3 part of the lamina ; tendrils as long to half as long as the leaves. Leaves of the climbing stems scattered, coriaceous, lanceolate, 15 to 35 cm long the petiole inclusive, 4 to 6 cm broad, acute or shortly acuminate, usually attenuate into the narrowly winged petiole, which is at most 10 cm long and dilated at the base to a semiamplexicaul sheath ; pennate nerves numerous, parallel and running almost straightly to the margin, the longitudinal nerves mostly 5, running parallel in the outer half of the lamina ; tendrils as long as to 1 1/2 times as long as the leaf, those with infundibuliform pitchers always with curl, the others without curl. Pitchers of the rosettes not known in the typical form, as far as known like those of the short stems. Pitchers of the short shoots and lower ones of the climbing stem obliquely urceolate, shortly incurved and abruptly widened from the hanging tendril, mostly 5 to 12 cm high, the lid side strongly curved, the other side straight with 2 fringed wings over the whole length, the wings 10 to 15 mm broad, the fringe segments 3 to 10 mm long, 1/2 to 1 mm apart ; mouth horizontal in front, abruptly incurved towards the lid, the peristome elongated into a short neck rarely longer than 1 cm, the outer margin involute, the inner part flat, up to 10 mm broad, the ribs 1/2 to 1/3 mm apart, the teeth of the interior margin about 3 to 5 times as long as broad ; inner surface of the pitcher wholly glandular, with minute overarched glands, about 1800 to 2000 on 1 cm2 ; lid elliptic-oblong, rarely slightly cordate, rounded or emarginate at the apex, 2 1/2 to 5 cm long, 1 1/2 to 2.8 cm broad, the lower surface with numerous not deepened glands, without appendage or keel ; spur 5 to 15 mm long, not branched, flattened, inserted close to the lid. Upper pitchers gradually or abruptly originating from the hanging end of the tendril, with a curve 15 to 20 mm wide, infundibuliform, about 8 to 13 cm high, 3 to 4 cm wide, slightly contracted under the mouth, mostly with 2 prominent ribs, rarely with 2 fringed wings, the wings up to 2 mm broad, the fringe up to 8 mm long ; mouth oblique, elevated towards the lid into a short neck mostly 1 cm high, the outer margin involute, the inner part flat, 10 to 15 cm broad, the ribs 1/2 to l/3 mm apart, the teeth of the inner margin about 3 times as long as broad ; inner surface of the pitcher wholly glandular with overarched glands, lid elliptic-oblong, rounded or emarginate at the apex, rounded at the base, with few large and many small not deepened glands on the under surface ; spur about 10 mm long, flattened, not branched. Male inflorescence a cylindrical raceme, the peduncle shorter than the raceme ; most pedicels 2-flowered, with a filiform bract above the base or not, 15 to 20 mm long, the upper ones 1-flowered. Tepals elliptical or broadly elliptical, rounded at the apex, about 5 to 7 mm long, the staminal column about 5 mm long, the anthers included, which are situated in 1 series and apical group. Female inflorescence almost like the male one, the lower pedicels sometimes 3-flowered. Ovary sessile. Fruit slender, 20 to 25 mm long, the valves lanceolate, strongly attenuate towards both ends, about 2 1/2 to 3 mm broad in the middle. Seeds filiform, about 10 to 15 mm long, the nucleus not wrinkled, delicately prickly. Indumentum densely tomentose in all parts when young, composed of intricate stellate hairs, later partly persistent or wholly deciduous on the stems and the petioles, sparse on the leaves beneath and on the pitchers, mixed with longer hairs and persistent on the inflorescences and the outer surface of the perigone, the staminal column glabrous from the beginning, the ovary densely tomentose, the fruit glabrescent, at length with brown points only. Colour in herbarium specimens: the leaves fallow-dun above, other parts red-brown. (Description after all the plants seen by the author.)

MALAY PENINSULA. Singapore: Jurong Reserve, 10 I 1889, H. S. (f) ; Chan Chu Kang, 15 X 1889, GOODENOUGH 1601, H. S. (m, f).

SUMATRA. Res. Eastcoast: Asahan, Tandjong Pasir, 60 m, 15 III 1925, YATES 1394, H. U. C. (0) ; H. B. (0) ; Res. Bangka: TEYSMANN, H. A. R. T. 000257 & 000258, vern. name: ketakong babie ; TEYSMANN 3512 & 3516, H. B. (0), vern. name: ketakong babie ; between Pelangas & Djeboes, TEYSMANN, H. B. (0), vern. name: ketakong babie.

BORNEO. British North Borneo: IX 1916-II 1917, VILLAMIL 261 bis, H. B. (0) ; Sarawak: native coll. 1790, H. B. (f) ; Res. Western Division: upper course of the Sg. Kapoeas, 1896-1897, JAHERI (Exp. NIEUWENHUIS) 63, H. B. (f) ; Res. Southern & Eastern Division: probably southern coast, 1859-1860, DE VRIESE & TEYSMANN, H. L. B. 908,129-1821 & -1822 (0).

The synonymy of N. Hookeriana has been very much confused, till in 1908 MACFARLANE has elucidated it. The principal cause of the confusion was the fact, that N. Hookeriana was not distinguished from N. Rafflesiana or considered as a variety of this species. MACFARLANE supposes N. Hookeriana to be a hybrid of N. ampullaria and N. Rafflesiana. I can quite join that opinion, since N. Hookeriana in a very striking manner is intermediate between the two species, and there is no argument against this supposition. In the herbaria we often find N. Hookeriana intermixed with N. ampullaria or N. Rafflesiana, and this proves that it at least often grows along with them. The area of dispersion of N. Rafflesiana only little differs from the western part of the area of N. ampullaria, and N. Hookeriana has almost only been found in the regions in which both species occur. Therefore it is a pity, that the hybrid between the often cultivated N. ampullaria and N. Rafflesiana has never been made experimentally. A simple experiment could solve this question.

Vernacular names. In Bangka, Malay, ketakong babi, pig's ketakong ; see for the explanation of this name under N. gracilis.


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